Local endemism within the Western Ghats-sri Lanka biodiversity hotspot.
Bossuyt F; Meegaskumbura M; Beenaerts N; Gower DJ; Pethiyagoda R; Roelants K; Mannaert A; Wilkinson M; Bahir MM; Manamendra-Arachchi K; Ng PK; Schneider CJ; Oommen OV; Milinkovitch MC; Science (New York, N.Y.), 2004, vol. 306, issue 5695, p 479, ISSN 10959203. ISBN 10959203.

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